When lactose is available but not glucose, then some lactose enters the cell using pre-existing transport protein encoded by lacY. If this weren't the case, there would be no lacY transporter protein in the cellular membrane; consequently, the lac operon would not be able to detect the presence of lactose. The repressor protein is always expressed, but the lac operon (i.e. enzymes and transport proteins) are almost completely repressed, allowing for a small level of background expression. In the absence of lactose, the lac repressor, encoded by lacI, halts production of the enzymes and transport proteins encoded by the lac operon. It would be wasteful to produce enzymes when no lactose is available or if a preferable energy source such as glucose were available. LacY encodes β-galactoside permease, a membrane protein which becomes embedded in the Plasma membrane to enable the cellular transport of lactose into the cell. This lactose metabolism system was used by François Jacob and Jacques Monod to determine how a biological cell knows which enzyme to synthesize. It is often discussed in introductory molecular and cellular biology classes for this reason.
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